Publications
(* Student author; ** Post-doc author)

Summary from Google Scholar

2025
Steinthorsdottir M, Montañez IP, Royer DL, Mills BJW, Hönisch B. 2024. Phanerozoic atmospheric CO₂ reconstructed with proxies and models: current understanding and future directions. Geochemistry Treatise (3rd Ed.), 5: 467-492 (invited contribution).
   [reprint]

2024
Lowe AJ, Flynn AG, Butrim MJ, Baumgartner AB, Royer DL, Peppe DJ. 2024. Reconstructing terrestrial paleoclimate and paleoecology with fossil leaves using digital leaf physiognomy and leaf mass per area. Journal of Visualized Experiments, 212: e66838.
   [reprint and supplemental information]
   [video tutorial]

Torsvik TH, Royer DL, Marcilly CM, Werner SC. 2024. User-friendly carbon-cycle modelling and aspects of Phanerozoic climate change. Applied Computing and Geosciences, 23: 100180.
   [reprint]
   [supplemental information]
      User-friendly interface for GEOCARBSULFvolc long-term carbon cycle model

Zhang X**, Royer DL, Shi G, Ichinnorov N, Herendeen PS, Crane PR, Herrera F. 2024. Estimates of late Early Cretaceous CO₂ from Mongolia based on stomatal and isotopic analysis of Pseudotorellia. American Journal of Botany, 111: e16376.
   [reprint]
   [supplemental information]

2023
Hönisch B, Royer DL, Breecker DO, Bowen GJ, Polissar PJ, Ridgwell A, and 78 other co-authors of the Cenozoic CO₂ Proxy Integration Project (CenCO₂PIP) Consortium. 2023. Toward a Cenozoic history of atmospheric CO₂. Science, 382: eadi5177.
   [reprint]
   [supplemental information]
      Companion website that compiles and integrates Phanerozoic CO₂ estimates from proxies

2022
Royer DL, Hren MT. 2022. Bulk carbon isotopic variability within leaves. PALAIOS, 37: 411-417 (with cover).
   [reprint]
   [supplemental information]

Liang J, Leng Q, Xiao L, Höfig DF, Royer DL, Zhang YG, Yang H. 2022. Early Miocene redwood fossils from Inner Mongolia: CO₂ reconstructions and paleoclimate effects of a low Mongolian plateau. Review of Palaeobotany and Paylnology, 305: 104743.
   [reprint]
   [supplemental information]

Butrim MJ*, Royer DL, Miller IM, Dechesne M, Neu-Yagle N, Lyson TR, Johnson KR, Barclay RS. 2022. No consistent shift in leaf dry mass per area across the Cretaceous-Paleogene boundary. Frontiers in Plant Science, 13: 894690.
   [reprint]
   [supplemental information]

Milligan JN, Flynn AG, Kowalczyk JB, Barclay RS, Geng J, Royer DL, Peppe DJ. 2022. Moderate to elevated atmospheric CO₂ during the early Paleocene recorded by Platanites leaves of the San Juan Basin, New Mexico. Paleoceanography and Paleoclimatology, 37: e2021PA004408.
   [reprint]
   [supplemental information]

Liang J, Leng Q, Höfig DF, Niu G, Wang L, Royer DL, Burke K, Xiao L, Zhang YG, Yang H. 2022. Constraining conifer physiological parameters in leaf gas-exchange models for ancient CO₂ reconstruction. Global and Planetary Change, 209: 103737.
   [reprint]
   [supplemental information]

2021
Schrader J, Shi P, Royer DL, Peppe DJ, Gallagher RV, Li Y-R, Wang R, Wright IJ. 2021. Leaf size estimation based on leaf length, width and shape. Annals of Botany, 128: 395-406.
   [reprint]
   [supplemental information]
   [commentary in Annals of Botany]

Marcilly CM, Torsvik TH, Domeier M, Royer DL. 2021. New paleogeographic and degassing parameters for long-term carbon cycle models. Gondwana Research, 97: 176-203.
   [reprint]
   [supplemental information]

Torsvik TH, Svensen HH, Steinberger B, Royer DL, Jerram DA, Jones MT, Domeier M. 2021. Connecting the deep Earth and the atmosphere. In: Marquardt H, Ballmer M, Cottaar S, Konter J (Editors), Mantle Convection and Surface Expressions. Geophysical Monograph, 263: 413-453.
   [reprint]

Wong TE, Cui Y, Royer DL, Keller K. 2021. A tighter constraint on Earth-system sensitivity from long-term temperature and carbon-cycle observations. Nature Communications, 12: 3173.
   [reprint]
   [supplemental information]

Konrad W, Royer DL, Franks PJ, Roth-Nebelsick A. 2021. Quantitative critique of leaf-based paleo-CO₂ proxies: consequences for their reliability and applicability. Geological Journal, 56: 886-902.
   [reprint]

2020
Butrim MJ*, Royer DL. 2020. Leaf-economic strategies across the Eocene-Oligocene transition correlate with dry season precipitation and paleoelevation. American Journal of Botany, 107: 1772-1785.
   [reprint]
   [supplemental information]

2019
Breecker DO, Royer DL. 2019. The pedogenic formation of coal balls by CO₂ degassing through the rootlets of arborescent lycopsids. American Journal of Science, 319: 529-548.
   [reprint]

Hollis CJ and 40 co-authors. 2019. The DeepMIP contribution to PMIP4: methodologies for selection, compilation and analysis of latest Paleocene and early Eocene climate proxy data, incorporating version 0.1 of the DeepMIP database. Geoscientific Model Development, 12: 3149-3206.
   [reprint]
   [supplemental information]

McKee ML*, Royer DL, Poulos H. 2019. Experimental evidence for species-dependent responses in leaf shape to temperature: implications for paleoclimate inference. PLoS ONE, 14: e021884.
   [reprint]
   [supplemental information]

Royer DL, Moynihan KM*, McKee ML*, Londoño L, Franks PJ. 2019. Sensitivity of a leaf gas-exchange model for estimating paleoatmospheric CO₂ concentration. Climate of the Past, 15: 795-809.
   [reprint]
   [supplemental information]

Milligan JN*, Royer DL, Franks PJ, Upchurch GR, McKee ML*. 2019. No evidence for a large atmospheric CO₂ spike across the Cretaceous-Paleogene boundary. Geophysical Research Letters, 46: 3462-3472.
   [reprint]
   [supplemental information]

Hagen ER*, Royer DL, Moye RA*, Johnson KR. 2019. No large bias within species between the reconstructed areas of complete and fragmented fossil leaves. Palaios, 34: 43-48.
   [reprint]
   [supplemental information]

2018
Kowalczyk JB*, Royer DL, Miller IM, Anderson CW, Beerling DJ, Franks PJ, Grein M, Konrad W, Roth-Nebelsick A, Bowring SA, Johnson KR, Ramezani J. 2018. Multiple proxy estimates of atmospheric CO₂ from an early Paleocene rainforest. Paleoceanography and Paleoclimatology, 33: 1427-1438.
   [reprint]
   [supplemental information]
      R code and input file to run Franks' v2 model for estimating CO₂ from stomatal geometry and leaf carbon isotopic composition (data sets S7-S8 in supplement)
      Mathematica and Maple code to run Konrad's v2 model for estimating CO₂ from stomatal geometry and leaf carbon isotopic composition (data sets S12-S14 in supplement)
      R code and input file to run Fletcher's BRYOCARB v2 model for estimating CO₂ from liverwort leaf carbon isotopic composition (data sets S9-S11 in supplement)

Londoño L, Royer DL, Jaramillo C, Escobar J, Foster DA, Cárdenas-Rozo AL, Wood A. 2018. Early Miocene CO₂ estimates from a Neotropical fossil assemblage exceed 400 ppm. American Journal of Botany, 105: 1929-1937.
   [reprint]
   [supplemental information]

Royer DL, Moynihan KM*, Ariori C*, Bodkin G*, Doria G*, Enright K*, Hatfield-Gardner R*, Kravet E*, Nuttle CM*, Shepard L*, Ku TCW, O'Connell S, Resor PG. 2018. Tank bromeliads capture Saharan dust in El Yunque National Forest, Puerto Rico. Atmospheric Environment, 173: 325-329.
   [reprint]
   [supplemental information]

2017
Edwards CT, Saltzman MR, Royer DL, Fike DA. 2017. Oxygenation as a driver of the Great Ordovician Biodiversification Event. Nature Geoscience, 10: 925-929.
   [reprint]
   [supplemental information]
   [News and Views in Nature Geoscience]
   [press release]

Wolfe AP, Reyes AV, Royer DL, Greenwood DR, Doria G*, Gagen MH, Siver PA, Westgate JA. 2017. Middle Eocene CO₂ and climate reconstructed from the sediment fill of a subarctic kimberlite maar. Geology, 45: 619-622.
   [reprint]
   [supplemental information]

O'Connell S, Gilmore M, Johnson-Thornton R, Ku T, Patton P, Resor P, Royer D. 2017. Diverse students can be attracted to geoscience. GSA Today, 27(5): 76-77.
   [reprint]

Royer DL, Hren HT. 2017. Carbon isotopic fractionation between whole leaves and cuticle. Palaios, 32: 199-205.
   [reprint]
   [supplemental data file 1] [supplemental data file 2]

Foster GL, Royer DL, Lunt DJ. 2017. Future climate forcing potentially without precedent in the geological record. Nature Communications, 8: 14845.
   [reprint]
   [supplemental information]
   [commentary in New Scientist] [commentary in The Guardian] [commentary in The Conversation] [commentary in Climate Central]
   [press release]

Franks PJ, Royer DL. 2017. Comment on "Was atmospheric CO₂ capped at 1000 ppm over the past 300 million years?" by McElwain J. C. et al. [Palaeogeogr. Palaeoclimatol. Palaeoecol. 441 (2016) 653-658]. Palaeogeography, Palaeoclimatology, Palaeoecology, 472: 256-259.
   [reprint]

2016
Royer DL. 2016. Climate sensitivity in the geologic past. Annual Review of Earth and Planetary Sciences, 44: 277-293.
   [reprint]

2015
Peppe DJ, Royer DL. 2015. Can climate feel the pressure? Science, 348: 1210-1211.
   [reprint]

Zanne AE, Tank DC, Cornwell WK, Eastman JM, Smith SA, FitzJohn RG, McGlinn DJ, O'Meara BC, Moles AT, Reich PB, Royer DL, Soltis DE, Stevens PF, Westoby M, Wright IJ, Aarssen L, Bertin RI, Calaminus A, Govaerts R, Hemmings F, Leishman MR, Oleksyn J, Soltis PS, Swenson NG, Warman L, Beaulieu JM. 2015. Reply to "Doubtful pathways to cold tolerance in plants". Nature, 521: doi:10.1038/nature14394.
   [reprint]

2014
Maxbauer DP*, Royer DL, LePage BA. 2014. High Arctic forests during the middle Eocene supported by moderate levels of atmospheric CO₂. Geology, 42: 1027-1030.
   [reprint]
   [supplemental information]

Royer DL, Donnadieu Y, Park J, Kowalczyk J*, Goddéris Y. 2014. Error analysis of CO₂ and O₂ estimates from the long-term geochemical model GEOCARBSULF. American Journal of Science, 314: 1259-1283 (with cover).
   [reprint]
   [R code and input files to run Berner's GEOCARBSULFvolc CO₂ model]

Blonder B, Royer DL, Johnson KR, Miller I, Enquist BJ. 2014. Plant ecological strategies shift across the Cretaceous-Paleogene boundary. PLoS Biology, 12(9): e1001949.
   [reprint]
   [supplemental information]
   [commentary in PLoS Biology] [commentary in Newsweek]
   [press release]

Franks PJ, Royer DL, Beerling DJ, Van de Water PK, Cantrill DJ, Barbour MM, Berry JA. 2014. New constraints on atmospheric CO₂ concentration for the Phanerozoic. Geophysical Research Letters, 41: 4685-4694.
   [reprint]
   [supplemental information]
      R code and input file to run Franks' v1 model for estimating CO₂ from stomatal geometry and leaf carbon isotopic composition
   [commentary in Science]

Cornwell WK, Westoby M, Falster DS, FitzJohn RG, O'Meara BC, Pennell MW, McGlinn DJ, Eastman JM, Moles AT, Reich PB, Tank DC, Wright IJ, Aarssen L, Beaulieu JM, Kooyman RM, Leishman MR, Miller ET, Niinemets Ü, Oleksyn J, Ordonez A, Royer DL, Smith SA, Stevens PF, Warman L, Wilf P, Zanne AE. 2014. Functional distinctiveness of major plant lineages. Journal of Ecology, 102: 345-356 (with cover).
   [reprint]
   [supplemental information]

Peppe DJ, Lemons CR, Royer DL, Wing SL, Wright IJ, Lusk CH, Rhoden CH*. 2014. Biomechanical and leaf-climate relationships: a comparison of ferns and seed plants. American Journal of Botany, 101: 338-347.
   [reprint]

Zanne AE, Tank DC, Cornwell WK, Eastman JM, Smith SA, FitzJohn RG, McGlinn DJ, O'Meara BC, Moles AT, Reich PB, Royer DL, Soltis DE, Stevens PF, Westoby M, Wright IJ, Aarssen L, Bertin RI, Calaminus A, Govaerts R, Hemmings F, Leishman MR, Oleksyn J, Soltis PS, Swenson NG, Warman L, Beaulieu JM. 2014. Three keys to the radiation of angiosperms into freezing environments. Nature, 506: 89-92.
   [reprint] [corrigendum]
   [supplemental information]

Royer DL. 2014. Atmospheric CO₂ and O₂ during the Phanerozoic: tools, patterns, and impacts. Geochemistry Treatise (Second Edition), 6: 251-267 (invited contribution).
   [reprint]

2013
Royer DL, Chernoff B. 2013. Diversity in neotropical wet forests during the Cenozoic linked more to atmospheric CO₂ than temperature. Proceedings of the Royal Society B, 280: 20131024.
   [reprint]

2012
Rohling EJ, Sluijs A, Dijkstra HA, Köhler P, van de Wal RSW, von der Heydt AS, Beerling D, Berger A, Bijl PK, Crucifix M, DeConto R, Drijfhout SS, Fedorov A, Foster G, Ganopolski A, Hansen J, Hönisch B, Hooghiemstra H, Huber M, Huybers P, Knutti R, Lea DW, Lourens LJ, Lunt D, Masson-Demotte V, Medina-Elizalde M, Otto-Bliesner B, Pagani M, Pälike H, Renssen H, Royer DL, Siddall M, Valdes P, Zachos JC, Zeebe RE. 2012. Making sense of palaeoclimate sensitivity. Nature, 491: 683-691.
   [reprint]
   [supplemental information]

Royer DL. 2012. Leaf shape responds to temperature but not CO₂ in Acer rubrum. PLoS ONE, 7(11): e49559.
   [reprint]

Royer DL. 2012. Climate reconstruction from leaf size and shape: new developments and challenges. In: Ivany LC, Huber BT (eds). Reconstructing Earth's Deep-Time Climate—The State of the Art in 2012. Paleontological Society Papers, 18: 195-212 (invited contribution).
   [reprint]

Poulos H, Bannon B, Isard J, Stonebraker P, Royer D, Yohe G, Chernoff B. 2012. The world through an interdisciplinary lens. Academe, 98(5): 41-43.
   [reprint] [web version]

Royer DL, Pagani M, Beerling DJ. 2012. Geobiological constraints on Earth system sensitivity to CO₂ during the Cretaceous and Cenozoic. Geobiology, 10: 298-310.
   [reprint]

Royer DL, Peppe DJ, Wheeler EA, Niinemets Ü. 2012. Roles of climate and functional traits in controlling toothed vs. untoothed leaf margins. American Journal of Botany, 99: 915-922.
   [reprint]
   [supplemental information]

Hönisch B, Ridgwell A, Schmidt DN, Thomas E, Gibbs SJ, Sluijs A, Zeebe R, Kump L, Martindale RC, Greene SE, Kiessling W, Ries J, Zachos JC, Royer DL, Barker S, Marchitto TM, Moyer R, Pelejero C, Ziveri P, Foster GL, Williams B. 2012. The geological record of ocean acidification. Science, 335: 1058-1063.
   [reprint]
   [supplemental information]

2011
Nicotra AB, Leigh A, Boyce CK, Jones CS, Niklas KJ, Royer DL, Tsukaya H. 2011. The evolution and functional significance of leaf shape in the angiosperms. Functional Plant Biology, 38: 535-552 (with cover).
   [reprint]

Beerling DJ, Royer DL. 2011. Convergent Cenozoic CO₂ history. Nature Geoscience, 4: 418-420.
   [reprint]
   [supplemental information]
   [editorial]

Doria G*, Royer DL, Wolfe AP, Fox A, Westgate JA, Beerling DJ. 2011. Declining atmospheric CO₂ during the late Middle Eocene climate transition. American Journal of Science, 311: 63-75.
   [reprint]

Park J, Royer DL. 2011. Geologic constraints on the glacial amplification of Phanerozoic climate sensitivity. American Journal of Science, 311: 1-26.
   [reprint]
   [commentary in Mother Jones]

Peppe DJ**, Royer DL, Cariglino B*, Oliver SY*, Newman S*, Leight E*, Enikolopov G*, Fernandez-Burgos M*, Herrera F, Adams JM, Correa E, Currano ED, Erickson JM, Hinojosa LF, Iglesias A, Jaramillo CA, Johnson KR, Jordan GJ, Kraft N, Lovelock EC, Lusk CH, Niinemets Ü, Peñuelas J, Rapson G, Wing SL, Wright IJ. 2011. Sensitivity of leaf size and shape to climate: global patterns and paleoclimatic applications. New Phytologist, 190: 724-739.
   [reprint]
   [supplemental information]
   [original leaf images]
   [commentary by Burnham & Tonkovich]
   [press release]

2010
Peppe DJ**, Royer DL, Wilf P, Kowalski EA. 2010. Quantification of large uncertainties in fossil leaf paleoaltimetry. Tectonics, 29: TC3015.
   [reprint]
   [supplemental information]

Royer DL, Miller IM, Peppe DJ**, Hickey LJ. 2010. Leaf economic traits from fossils support a weedy habit for early angiosperms. American Journal of Botany, 97: 438-445.
   [reprint]
   [supplemental information]
   [press release]

Royer DL. 2010. Fossil soils constrain ancient climate sensitivity. Proceedings of the National Academy of Sciences USA, 107: 517-518. (invited commentary)
   [reprint]

2009
Royer DL, Meyerson LA, Robertson KM, Adams JM. 2009. Phenotypic plasticity of leaf shape along a temperature gradient in Acer rubrum. PLoS ONE, 4(10): e7653.
   [reprint]
   [supplemental information]

Royer DL, Kooyman RM, Little SA, Wilf P. 2009. Ecology of leaf teeth: A multi-site analysis from an Australian subtropical rainforest. American Journal of Botany, 96: 738-750.
   [reprint]
   [supplemental Appendix S1] [supplemental Appendix S2]

2008
Hansen J, Sato M, Kharecha P, Beerling D, Berner R, Masson-Delmotte V, Pagani M, Raymo M, Royer DL, Zachos JC. 2008. Target atmospheric CO₂: where should humanity aim? Open Atmospheric Science Journal, 2: 217-231.
   [reprint]
   [supplemental information]
   [commentary in The Guardian] [commentary in The New York Times] [science behind 350.org]

Royer DL. 2008. Nutrient turnover rates in ancient terrestrial ecosystems. Palaios, 23: 421-423. (invited "Spotlight")
   [reprint]

Royer DL, McElwain JC, Adams JM, Wilf P. 2008. Sensitivity of leaf size and shape to climate within Acer rubrum and Quercus kelloggii. New Phytologist, 179: 808-817.
   [reprint]
   [supplemental information]
   [original leaf images]

Currano ED, Wilf P, Wing SL, Labandeira CC, Lovelock EC, Royer DL. 2008. Sharply increased insect herbivory during the Paleocene-Eocene Thermal Maximum. Proceedings of the National Academy of Sciences USA, 105: 1960-1964.
   [reprint]
   [supplemental information]
   [commentary by DeLucia et al.] [commentary in Nature Reports Climate Change] [commentary in Nature Geoscience]

Royer DL. 2008. Linkages between CO₂, climate, and evolution in deep time. Proceedings of the National Academy of Sciences USA, 105: 407-408. (invited commentary)
   [reprint]

2007
Royer DL, Sack L, Wilf P, Lusk CH, Jordan GJ, Niinemets Ü, Wright IJ, Westoby M, Cariglino B, Coley PD, Cutter AD, Johnson KR, Labandeira CC, Moles AT, Palmer MB, Valladares F. 2007. Fossil leaf economics quantified: calibration, Eocene case study, and implications. Paleobiology, 33: 574-589.
   [reprint]
   [online appendix A] [online appendix B]
   [Editors' Choice in Science]

Royer DL, Berner RA, Park J. 2007. Climate sensitivity constrained by CO₂ concentrations over the past 420 million years. Nature, 446: 530-532.
   [reprint]
   [supplemental information]
   [commentary in The Christian Science Monitor] [commentary in environmentalresearchweb] [commentary in ars technica]
   [podcast at Nature] [interview with Where We Live (WNPR)]
   [press release]

2006
Royer DL. 2006. CO₂-forced climate thresholds during the Phanerozoic. Geochimica et Cosmochimica Acta, 70: 5665-5675. (invited contribution, with cover)
   [reprint]
   [supplemental information]

Royer DL, Wilf P. 2006. Why do toothed leaves correlate with cold climates? Gas-exchange at leaf margins provides new insights into a classic paleotemperature proxy. International Journal of Plant Sciences, 167: 11-18.
   [reprint]
   [commentary in The Union] [commentary in Discovery News]

2005
Fletcher BJ, Beerling DJ, Brentnall SJ, Royer DL. 2005. Fossil bryophytes as recorders of ancient CO₂ levels: Experimental evidence and a Cretaceous case study. Global Biogeochemical Cycles, 19: GB3012.
   [reprint]

Royer DL, Wilf P, Janesko DA*, Kowalski EA, Dilcher DL. 2005. Correlating climate and plant ecology to leaf size and shape: potential proxies for the fossil record. American Journal of Botany, 92: 1141-1151.
   [reprint]
   [supplemental information]
   [original leaf images]

Royer DL, Osborne CP, Beerling DJ. 2005. Contrasting seasonal patterns of carbon gain in evergreen and deciduous trees of ancient polar forests. Paleobiology, 31: 141-150.
   [reprint]

2004
Osborne CP, Royer DL, Beerling DJ. 2004. Adaptive role of leaf habit in extinct polar forests. International Forestry Review, 6: 181-186.
   [reprint]

Royer DL, Berner RA, Montañez IP, Tabor NJ, Beerling DJ. 2004. CO₂ as a primary driver of Phanerozoic climate change: Reply. GSA Today, 14(7): 18.
   [reprint]

Royer DL, Berner RA, Montañez IP, Tabor NJ, Beerling DJ. 2004. CO₂ as a primary driver of Phanerozoic climate change. GSA Today, 14(3): 4-10.
   [reprint]
   [supplemental information]
   [commentary in Science] [commentary in The New York Times] [commentary in Natural History]
   [press release]

2003
Royer DL, Osborne CP, Beerling DJ. 2003. Carbon loss by deciduous trees in a CO₂-rich ancient polar environment. Nature, 424: 60-62.
   [reprint]
   [commentary in BBC News]
   [press release]

Royer DL. 2003. Estimating latest Cretaceous and Tertiary atmospheric CO₂ from stomatal indices. In: Wing SL, Gingerich PD, Schmitz B, Thomas E (eds). Causes and Consequences of Globally Warm Climates in the Early Paleogene. Geological Society of America Special Paper, 369: 79-93.
   [reprint]

Kauffman SJ, Royer DL, Chang S, Berner RA. 2003. Export of chloride after clear-cutting in the Hubbard Brook sandbox experiment. Biogeochemistry, 63: 23-33.
   [reprint]

Royer DL, Hickey LJ, Wing SL. 2003. Ecological conservatism in the “living fossil” Ginkgo. Paleobiology, 29: 84-104.
   [reprint]

2002
Beerling DJ, Lake JA, Berner RA, Hickey LJ, Taylor DW, Royer DL. 2002. Carbon isotope evidence implying high O₂/CO₂ ratios in the Permo-Carboniferous atmosphere. Geochimica et Cosmochimica Acta, 66: 3757-3767.
   [reprint]

Royer DL, Osborne CP, Beerling DJ. 2002. High CO₂ increases the freezing sensitivity of plants: implications for paleoclimatic reconstructions from fossil floras. Geology, 30: 963-966.
   [reprint]

Beerling DJ, Lomax BH, Royer DL, Upchurch GR, Kump LR. 2002. An atmospheric pCO₂ reconstruction across the Cretaceous-Tertiary boundary from leaf megafossils. Proceedings of the National Academy of Sciences USA, 99: 7836-7840.
   [reprint]
   [commentary in New Scientist] [commentary in National Geographic] [commentary in BBC News]

Beerling DJ, Royer DL. 2002. Fossil plants as indicators of the Phanerozoic global carbon cycle. Annual Review of Earth and Planetary Sciences, 30: 527-556.
   [reprint]

Beerling DJ, Royer DL. 2002. Reading a CO₂ signal from fossil stomata. New Phytologist, 153: 387-397.
   [reprint]

Royer DL. 2002. Estimating latest Cretaceous and Tertiary PCO₂ from stomatal indices [Ph.D. thesis]. Yale University, New Haven, 163 p.
   [reprint]

2001
Royer DL, Wing SL, Beerling DJ, Jolley DW, Koch PL, Hickey LH, Berner RA. 2001. Paleobotanical evidence for near present day levels of atmospheric CO₂ during part of the Tertiary. Science, 292: 2310-2313.
   [reprint]
   [commentary in Geology Today]

Royer DL, Berner RA, Beerling DJ. 2001. Phanerozoic atmospheric CO₂ change: Evaluating geochemical and paleobiological approaches. Earth-Science Reviews, 54: 349-392.
   [reprint]
   [Editors' Choice in Science]

Royer DL. 2001. Stomatal density and stomatal index as indicators of paleoatmospheric CO₂ concentration. Review of Palaeobotany and Palynology, 114: 1-28.
   [reprint]

2000
Gough L, Shaver GR, Carroll J, Royer DL, Laundre JA. 2000. Vascular plant species richness in Alaskan arctic tundra: The importance of soil pH. Journal of Ecology, 88: 54-66.
   [reprint]

Royer DL. 2000. Depth to pedogenic carbonate horizon as a paleoprecipitation indicator?: Reply. Geology, 28: 572-573.
   [reprint]

1999
Royer DL. 1999. Depth to pedogenic carbonate horizon as a paleoprecipitation indicator? Geology, 27: 1123-1126.
   [reprint]
   [supplemental information]